Static Mesh Only enabled Foliage Actor causes serialization error in packaged game

The crash can be worked around by modifying FFoliageISMActor::Serialize as following code :

void FFoliageISMActor::Serialize(FArchive& Ar)
{
#if WITH_EDITORONLY_DATA
    if (!Ar.IsCooking()) // add this if statement
    {
        Ar << Guid;
        ClientHandle.Serialize(Ar);
        Ar << ISMDefinition;
        Ar << ActorClass;
    }
#endif
}
 

1. Create a blueprint based on Actor
2. Add static mesh component and static mesh to the blueprint
3. Create a Actor Foliage asset
4. Open the asset
5. Assign the created blueprint to the actor foliage asset
6. Turn Static Mesh Only on
7. Save and close foliage actor window
8. Create a level with Basic template
9. Enter foliage mode
10. Add the actor foliage asset to the foliage panel
11. Paint (note : Need changing Floor Actor mobility to Static)
12. Save the level
13. Open project settings
14. Set GameDefaultMap and EditorStartMap to the created level
15. Create Package for Windows 
16. Run the package

The packaged game crushes with following error message :

ObjectSerializationError: /Game/P_ISMActorFoliage (0x560D969E43985BE3) /Game/P_ISMActorFoliage (0x560D969E43985BE3) - InstancedFoliageActor /Game/P_ISMActorFoliage.P_ISMActorFoliage:PersistentLevel.InstancedFoliageActor_0: Bad import index 373652847/67.

LogWindows: Error: [Callstack] 0x00007ff63ed2f202 UnrealGame.exe!FExportArchive::HandleBadImportIndex() [D:\dev\UnrealEngine-5.3.1\Engine\Source\Runtime\CoreUObject\Private\Serialization\AsyncLoading2.cpp:1709]
LogWindows: Error: [Callstack] 0x00007ff63ed1652b UnrealGame.exe!FExportArchive::operator<<() [D:\dev\UnrealEngine-5.3.1\Engine\Source\Runtime\CoreUObject\Private\Serialization\AsyncLoading2.cpp:1758]
LogWindows: Error: [Callstack] 0x00007ff63ed02d40 UnrealGame.exe!FArchiveUObject::SerializeObjectPtr() [D:\dev\UnrealEngine-5.3.1\Engine\Source\Runtime\CoreUObject\Private\Serialization\ArchiveUObject.cpp:90]
LogWindows: Error: [Callstack] 0x00007ff6429a3961 UnrealGame.exe!operator<<<TSetElement<TTuple<TObjectPtr<UFoliageType>,TUniqueObj<FFoliageInfo> > >,TSparseArrayAllocator<TSizedDefaultAllocator<32>,FDefaultBitArrayAllocator> >() [D:\dev\UnrealEngine-5.3.1\Engine\Source\Runtime\Core\Public\Containers\SparseArray.h:1496]
LogWindows: Error: [Callstack] 0x00007ff6429a9bac UnrealGame.exe!TSetPrivateFriend::Serialize<TTuple<TObjectPtr<UFoliageType>,TUniqueObj<FFoliageInfo> >,TDefaultMapHashableKeyFuncs<TObjectPtr<UFoliageType>,TUniqueObj<FFoliageInfo>,0>,FDefaultSetAllocator>() [D:\dev\UnrealEngine-5.3.1\Engine\Source\Runtime\Core\Public\Containers\Set.h:2201]
LogWindows: Error: [Callstack] 0x00007ff6429c279c UnrealGame.exe!AInstancedFoliageActor::Serialize() [D:\dev\UnrealEngine-5.3.1\Engine\Source\Runtime\Foliage\Private\InstancedFoliage.cpp:4390]
LogWindows: Error: [Callstack] 0x00007ff63ed24096 UnrealGame.exe!FAsyncPackage2::EventDrivenSerializeExport() [D:\dev\UnrealEngine-5.3.1\Engine\Source\Runtime\CoreUObject\Private\Serialization\AsyncLoading2.cpp:6370]
LogWindows: Error: [Callstack] 0x00007ff63ed26d82 UnrealGame.exe!FAsyncPackage2::Event_ProcessExportBundle() [D:\dev\UnrealEngine-5.3.1\Engine\Source\Runtime\CoreUObject\Private\Serialization\AsyncLoading2.cpp:5796]
LogWindows: Error: [Callstack] 0x00007ff63ed27a5d UnrealGame.exe!FEventLoadNode2::Execute() [D:\dev\UnrealEngine-5.3.1\Engine\Source\Runtime\CoreUObject\Private\Serialization\AsyncLoading2.cpp:4312]
LogWindows: Error: [Callstack] 0x00007ff63ed2805b UnrealGame.exe!FAsyncLoadEventQueue2::ExecuteSyncLoadEvents() [D:\dev\UnrealEngine-5.3.1\Engine\Source\Runtime\CoreUObject\Private\Serialization\AsyncLoading2.cpp:4500]
LogWindows: Error: [Callstack] 0x00007ff63ed41f06 UnrealGame.exe!FAsyncLoadingThread2::Run() [D:\dev\UnrealEngine-5.3.1\Engine\Source\Runtime\CoreUObject\Private\Serialization\AsyncLoading2.cpp:7662]
LogWindows: Error: [Callstack] 0x00007ff63e9c4132 UnrealGame.exe!FRunnableThreadWin::Run() [D:\dev\UnrealEngine-5.3.1\Engine\Source\Runtime\Core\Private\Windows\WindowsRunnableThread.cpp:149]
LogWindows: Error: [Callstack] 0x00007ff63e9b7f47 UnrealGame.exe!FRunnableThreadWin::GuardedRun() [D:\dev\UnrealEngine-5.3.1\Engine\Source\Runtime\Core\Private\Windows\WindowsRunnableThread.cpp:79]
LogWindows: Error: [Callstack] 0x00007ff849b77344 KERNEL32.DLL!UnknownFunction []

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