Crash/ensure when linking anim layers from animation update

Originally reported here: https://udn.unrealengine.com/questions/562047/linked-layer-animation-blueprint-proxy-missing-ske.html

When linking an anim layer during update, we get either an ensure or crash (missing skeleton) depending on the structure of the graph.

Ensure message: "PreUpdate isn't called. This could potentially cause other issues."

This seems like a valid code path to support. Seems we'd either need to queue the link (introducing a frame delay?) or do some sort of fixup.

We haven't encountered this internally because we've so far only been linking layers from gameplay code.

Call LinkAnimClassLayers() from NativeUpdateAnimation or BlueprintUpdateAnimation.

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