Looks like we are missing code in the Evaluate methods similar to what we are doing in 
DecompressPose to account for compatible skeletons.  The easiest way to repro is in a standalone game because that is currently using the raw transforms (a separate bug).
There's no existing public thread on this issue, so head over to Questions & Answers just mention UE-193895 in the post.
| 2 | 
| Component | UE - Anim - Runtime | 
|---|---|
| Target Fix | 5.4 | 
| Created | Aug 25, 2023 | 
|---|---|
| Resolved | Sep 13, 2023 | 
| Updated | Apr 17, 2024 |